Failed to invite user - Error 554

I'm getting the following error when trying to invite a user to join upsource (I have redacted the actual email address but it I am using a vaild one):

Failed to invite 1 user email@domain.com with error 554 Your message was received but it could not be saved. Please retry later

Anyone have any more details on what a 554 error is?

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It appears the 554 error is an SMTP error, so I'll investigate the mail server.

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